Systematic revision, cladistic analysis and biogeography of the genera Phloeotragus and Phloeopemon (Coleoptera, Anthribidae, Anthribinae, Ptychoderini)

The genera Phloeotragus Schoenherr, 1823 comprises thirteen Afrotropical species and Phloeopemon Schoenherr, 1839 with four species is primarily Oriental and occurring in Indo-Malaya. Only one species (Phloeopemon acuticornis) reaching the Japan in Paleartic region. A revision and cladistic analysis of the genera is conducted, including 117 characters from body vestiture, external morphology, wing venation, male and female terminalia and rectal loop. Twenty-four taxa were included in the cladistic analysis, seven of which as outgroups. The eighteen most parsimonious cladograms obtained and the strict consensus show that Phloeopemon is sister group of Phloeotragus supported on the basis of the eight tergite and first tarsomere longer than second, more so in hind legs. Phloeotragus is supported by dorsum of rostrum not carinate, anterior margin of pronotum strongly emarginated and pronotum with four isolated impressions on disk. These unique characters distinguish Phloeotragus from another genera. Phloeopemon is well supported by four uniquely derived character states: 1) a narrow vitta clothed with bright scales on pronotum; 2) femora with integument black spot - clothed by denser blackish-brown pubescence; 3) suberect setae on inner face of the mesotibiae; and 4) pronotum coarsely and confluent punctate. A key to species, illustrations and maps of distribution are provided. The biogeography based on patterns of distribution of the genera are discussed.