Eyeing the evolution of Odonate color vision

In insects, different parts of the eye and different cells in each ommatidium are capable of capturing different wavelengths of light.  Opsins are light sensitive proteins in the eye that can help organism discriminate color.  The ancestral insect eye is thought to have had at least three opsin copies: ultra violet, blue, and long wavelength.  However, current research has shown that insects have duplicated (and even lost) opsin copies throughout their evolution.  It is unclear how widely opsin gene duplications and losses are distributed throughout insects.  Dragonflies and damselflies (Odonata, a primitive winged insect group) are entirely visual organisms, yet little research has been conducted on the origin and function of their color vision system.  The current hypotheses are that dragonflies have as many as five opsins and damselflies as many as three.  Using transcriptome, PCR data, and physiological evidence gathered in the lab and from the literature the opsin genes of >15 species of odonates were examined in a phylogenetic framework to determine the following: 1) Are there more than five opsin copies in dragonflies and more than three in damselflies? 2) Are there opsin duplications and in what lineages do these duplications occur? 3) Are there any biological reasons for such duplications/losses? My research will provide a much-needed view of the evolution of vision within a basal insect group and provide a deeper context for the evolution of insect visual systems.