Richard Stouthamer, richard.stouthamer@ucr.edu, University of California - Riverside, Department of Entomology, Riverside, CA, James E Russell, james.russell@email.ucr.edu, University of California - Riverside, Department of Entomology, Riverside, CA, and Fabrice Vavre, vavre@biomserv.univ-lyon1.fr, Universite de Claude Bernarde-Lyon, Laboratoire de Biometrie et Biologie Evolutive, UMR CNRS 5558-Bat. G. Mendel, 16 rue Raphael Dubois, Villeurbanne, France.