The larvae of the family Sciomyzidae (Diptera: Acalyptratae) display a wide range of feeding behaviors, being predators, parasitoids, or saprophages of a wide variety of molluscs (mostly gastropods). It is an ideal group of organisms with which to study the evolution of feeding behavior and associated ecological and morphological characteristics within a phylogenetic framework. The genus Tetanocera is particularly interesting because its species occupy five of the 17 currently recognized sciomyzid feeding groups. This project has three specific OBJECTIVES: (1) Construct a robust estimate of phylogeny for Tetanocera, (2) Estimate the evolutionary transitions in larval feeding behaviors and habitats that have occurred during Tetanocera phylogenesis, and (3) Investigate potential correlations among larval ecological characteristics (e.g., aquatic vs. terrestrial habitat) and larval morphology. Eighteen species of Tetanocera and eight additional sciomyzid genera were used with a variety of phylogenetic algorithms to build the phylogeny and conduct the above investigations. It was found that lineages of Tetanocera left the aquatic habitat between 3-6 times independently during their evolutionary history. Such habitat shifts were accompanied by a suite of morphological changes, and their correlations were found to be statistically significant using phylogenetic comparative methods.