The odd staphylinid genus Arrowinus Bernhauer, 1935 is revised, with three new species and the first larval description. The flightless species show apparent relict distribution patterns in southern South Africa. The genus has been placed in the subtribe Quediina of Staphylininae: Staphylinini, but neither adult nor larval characters agree with that placement. A phylogenetic analysis of exemplars of all named tribes and subtribes of the subfamily Staphylininae plus four outgroups was conducted, by means of partitioned analyses of adults and larvae separately plus a combined analysis. Results of all analyses support the exclusion of Arrowinus from Quediina and from Staphylinini, and proposal of a new tribe of Staphylininae for it, as a basal phylogenetic relict within the subfamily. Implications of the analyses for phylogeny and classification of other Staphylininae are discussed.
Species 1: Coleoptera Staphylinidae Arrowinus phaenominalis
Keywords: partitioned analysis, flightlessness
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